When publishing mogul Bradford Meade (Alan Dale) hands the reins of Mode, the bible of the fashion industry, over to this son, Daniel (Eric Mabius), he specifically hires Betty Suarez as his son s new assistant - mostly because she's the only woman in NYC Daniel won't sleep with. Though this "player" is reluctant to accept her at first, Betty's indomitable spirit and bright ideas will eventually win him over. Neither of them really knows the ins and outs of the fashion world, but the two are a formidable team against those who will do anything to see them fall.

These are Caleb's (recurring guest star Alan Dale) last days as a bachelor, and he's spending them in Las Vegas. Seth and Ryan join him in "Sin City" for the celebration while a reluctant Kristen plans a bachelorette party for Julie back home in Newport Beach. Meanwhile, Marissa and Theresa (recurring guest star Navi Rawat) are learning more about each other, but what Marissa finds out about Teresa could have big consequences.

Having seriously dated for one week, Marissa and Ryan are progressing slowly. Summer (recurring guest star Rachel Bilson) tries to deny her true feelings for Seth as she witnesses Anna (recurring guest star Samaire Armstrong) and Seth getting closer. Meanwhile, the case against Kirsten's father, Caleb (recurring guest star Alan Dale), continues to cause friction between Sandy and Kirsten. And Julie (recurring guest star Melinda Clarke) tries to mend her relationship with Marissa, inviting her, Jimmy and Ryan to attend her charity event--where her new relationship is revealed.

Marissa and Ryan try to start over as friends, but it's not as easy as they had hoped. As Theresa (recurring guest star Navi Rawat), Ryan's ex-girlfriend from Chino, begins spending more time in Newport, Ryan discovers that she hasn't been honest about why she's there. Seth believes his relationship with Summer is great, but when she refuses to acknowledge him at school, he feels he has to teach her a lesson. Meanwhile, Sandy finds himself embroiled in another mess created by Caleb (recurring guest star Alan Dale). And Jimmy begins to suspect something is going on between Julie and Luke.

Marissa isn't ready to speak to Luke or her mother, Julie, following her discovery that the two have been having an affair. Luke is desperate to explain himself. When he gets into a near-fatal accident, however, both Marissa and Ryan decide to listen to what he has to say. Meanwhile, it's opening night for new restaurant The Lighthouse. Just prior to opening, Jimmy and Sandy are shocked to find out that their liquor license has been denied, and Sandy is faced with the choice of getting the license or keeping Jimmy as his partner. Also, Caleb (recurring guest star Alan Dale) uses the occasion to surprise everyone by proposing marriage to Julie.

Actors Vanessa Williams (Wilhelmina Slater), Alan Dale (Bradford Meade), Rebecca Romijn (Alexis Meade), and Eric Mabius (Daniel Meade) each convey what they really like about the show.

Michael Emerson and Alan Dale take us behind the scenes in London.

Catch a sneak peek at this season of "Ugly Betty."

America Ferrera and Salma Hayek discuss the message of "Ugly Betty."

Mark Consuelos is joining the Lindsay Lohan and Victoria Beckham Club by appearing on this season of "Ugly Betty."